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Meristics is an area of and which relates to counting quantitative features of animals and plants, such as the number of fins or scales in . A meristic (countable trait) can be used to describe a particular , or used to identify an unknown species. Meristic traits are often described in a shorthand notation called a meristic formula.

Meristic characters are the countable structures occurring in series (e.g. , , ). These characters are among the characters most commonly used for differentiation of species and . In the , scale counts have been most widely used for the differentiation of populations within species. In and the most notable differences among populations occur in counts of scales. Meristic comparison is used in and analysis.


Meristic analysis
A meristic study is often a difficult task. For example, counting the features of a fish is not as easy as it may appear. Many meristic analyses are performed on dead fish that have been preserved in alcohol. Meristic traits are less easily observed on living fish, though it is possible. On very small fish, a microscope may be required.

follow a basic set of rules when performing a meristic analysis, to remove as much ambiguity as possible. The specific practice, however, may vary depending on the type of fish. The methodology for counting meristic traits should be described by the specialist who performs the analysis.


Meristic formula
A meristic formula is a shorthand method of describing the way the bones (rays) of a 's are arranged. It is comparable to the floral formula for flowers.

Spine counts are given in Roman numerals, e.g. XI-XIV. Ray counts are given in Arabic numerals, e.g. 11–14.

The meristic formula of the ( Siganus luridus) is: D, XIV+10; A, VII+8-9; P, 16–17; V, I+3+I; GR, 18-22

This means the fish has 14 spiny rays (bones) in the first part of its (D), followed by 10 soft rays. A is the anal fin, P represents the (near the gills and eyes), V represents the or , and C is the caudal fin or tail (not indicated in this example). GR means gill raker count (see below).


Vertebral counts
The number of bones in the backbone is a feature which can also be used to classify fish species. Usually all the are counted. Vertebral counts may be split into abdominal (those associated with the body cavity) and caudal (tail) vertebrae. If there are sutures in the , components are counted, otherwise the urostyle is usually counted as one vertebra.


Gill raker counts
The number of gill rakers on the first can also be used to identify a fish species. Rakers are counted for the upper and lower limbs of the gill arch, and a raker at the joint of the upper and lower limbs is counted as of the lower. Counts for the upper and lower limbs are separated by a + sign and ranges are bracketed, e.g., GR: 3 + (4-5).


See also


External links
  • Chase, P. D. (2014) "Meristics". In: Steven X. Cadrin, Lisa A. Kerr and Stefano Mariani (Eds) Stock Identification Methods: Applications in Fishery Science, second edition, pp. 171–184. Academic Press. .
  • Fish identification using meristics Practical fishkeeping. 26 July 2010.

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